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Having walked to Stravithie along the B9131 (St Andrews to Anstruther), I took the unclassified road across to the A917, and walked back to St Andrews. (The milestones between St Andrews and Stravithie are described here.)
With the exception of the first one out of St Andrews the milestones on this stretch of road give the distances from St Andrews, Leuchars and Cupar, and Boarhills, Kingsbarns and Crail.
The photographs were taken on an increasingly cloudy afternoon at the end of November 2012.
Here is an account of the walk. (PDF file)
The unclassified road goes past Bonnytown and reaches the Coast road between Kinglassie and Boarhills Church, a mile or so from Boarhills itself.
The four-mile stone with the grass removed. The Milestones of Fife (2005) reports that the cap is missing from this stone. It also states that the distance from Boarhills should be given as half a mile rather than 1½, which would be consistent with the next stone. The stone is situated on the left-hand side of the road, going away from St Andrews, which is as stated in The Milestones of Fife, but the OS Landranger map marks it on the right-hand side.
The first milestone out of St Andrews is before the fork at Brownhills. It belongs to the same series as the stones on the Anstruther road, showing Newport instead of Cupar, and Anstruther, Kilrenny and Pittenweem instead of Boarhills, Kingsbarns and Crail. (It was photographed in bright morning sunshine on the walk to Stravithie.)